The Greek tortoise is also known as the spur-thighed tortoise for the large conical tubercle it has on each thigh. Several subspecies are recognised, which vary greatly in colour and size. The high, domed upper shell (carapace) is around 20 cm in length in T. g. graeca, but almost twice as large in T. g. ibera. The carapace varies from yellow or tan with black or dark-brown blotching to totally grey or black, while the lower shell (plastron) may be yellow to greenish-yellow, brown, or grey, with some dark-brown or black markings. Neck, limbs, and tail are yellowish brown to grey, while the head ranges from yellow to brown, grey, or black, with or without dark spotting. Large, overlapping scales cover the front of the forelimbs and there are five claws on each foot (2).